Adaptation and transformation have emerged as a key themes for human-environment research, especially in the context of rapid social-ecological changes. The 2008 global financial crisis constitutes a major driver of change with social-ecological ramifications that have yet to be fully explored. Using Greece, the poster child of the euro-crisis as a case-study, this dissertation examines how adaptive capacity is mobilized and even enhanced in times of crisis, paying particular attention to the role played by natural capital. To do so, I focus on the back-to-land trend whereby urbanites seek to engage in food production post-crisis (2008-onwards). In-depth qualitative analysis of back-to-landers’ motivations, experiences, and challenges is integrated with quantitative data about household demographics, incomes and assets, and land management characteristics. The dissertation is organized in three main result papers (chapters). The first seeks to understand why people turn to the land in times of crisis, and the role played by agency. The second analyzes the various assets that people mobilize in order to go back to the land, paying particular attention to the different mobilities necessary for their livelihood transformation. The third examines environmental safety nets in terms of material and non-material benefits that ecosystems provide to people. This research contributes to a wider social-ecological scholarship that seeks to understand how people adapt and transform when confronted with crises, focusing on how land and associated ecosystem services contribute to the resilience of these households, and the role played by agency in this process.